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I may be mistaken, but I don't think anyone at any weight has that many title defenses.
I'm sure someone will correct me if I'm wrong.
I gotta go with Ali. What he lacked in KO power (his only weakness in his prime imo), he made up for with his psychologic warfare and cerebral tactics. He was so good he's still the most recognizable face of all time in any sport... and last time I heard he was the most recognized person on the planet.
To me these things factor into the equation.
If we were going strictly on in ring criteria, then it would probably go to a lighter fighter like Sugar Ray Robinson possibly.