I don't understand the argument here - that the foot structure of deer, horses, dogs and cats evolved in a different manner than those of humans automatically equates to human beings being better at grappling than at running.
There's been considerable talk about human beings having the capacity to outrun horses and other prey animals over the course of a day.
http://www.nytimes.com/2009/10/27/health/27well.html
Persistence hunting - Wikipedia, the free encyclopedia
As of right now, it's a hypothesis that seems to have some science behind it as well as observation of certain aboriginal tribes (or tribes that have kept some traditions of distance running/hunting alive). It's not proven, but there's some slight rumblings that there could be scientific evidence enough behind it to get it to "theory" status someday. Theories are things like "the theory of evolution" or the "Big Bang theory". Lots of math, science, observations and brain power.
In terms of pure grappling, I think we humans lose to just about every large land mammal on earth and maybe a reptile/bird or two. Apes, big cats, bears, rhinos, big deer, cows, hyenas and so on. Crocodiles and cassowaries might be an even battle. We're not as fast, strong, stable, equipped with lethal instruments or fluid as these animals.
There's little to no science behind "human vs. animal x" because to test this stuff out would require a very large population of possibly insane humans willing to die or be severely wounded, a large captive population of big animals and some very questionable ethics and funding decisions.
However, the common sense test of considering this in a sort of rational way - to me - suggests that we humans are better at running in comparison to the animal kingdom than we are at grappling.
Whether we are closer to the "ideal" or the limits of peak human performance in running or in grappling is unknown to me and probably unstudied.
Right now, I know that Usain Bolt runs a hell of a lot faster than I do and ultra-marathon runners run a hell of a lot farther than I do. Buivasar Saitiev wrestles a hell of a lot better than I do and Rickson Gracie always wins by armbar or RNC.
However, I don't know if Usain Bolt is already the fastest human that will ever walk the earth, that Buivasar is the best human grappler in existence and so on. I don't know how far the human body can be taken and the Olympics/grappling sports/weird specialist sports are still experiencing progress and improvement in terms of human performance.
I don't know this stuff and while I'm somewhat intrigued by considering and finding out these things, FUCK this stupid bullshit that uses no scientific support, no common sense and serves only to support some kind of weird ass "bolstering" of submission grappling as a useful (and perhaps money-making) pursuit.
Fuck...